Andreas Lindén


Senior Researcher, PhD, Docent (Adjunct Professor)

Bioeconomy Research Team
Raseborgsvägen 9
FI-10600 Ekenäs


I am a theoretical ecologist with a wide range of interests within the field of population ecology and behavioural ecology. I am particularly dedicated to link empirical data to statistical models, which are tailored to provide explicit answers to ecologically relevant questions. I strive to jointly investigate practical ecological questions, develop ecological theory and methodology, as all these are linked to each other. Birds have been my focal study system, analysed in the light of various long-term data sets from monitoring programmes and birds observatories, but I am interested in other taxa and study systems as well.

I studied biology at the University of Helsinki (Finland), with ecology and evolutionary biology as my major subject. My PhD thesis was about statistical modelling of environmental effects in population dynamics, supervised by Esa Ranta and Veijo Kaitala. After this I worked three years as a post doctoral researcher at the University of Oslo, studying bird migration phenology and its population dynamical consequences, supervised by Nils Chr. Stenseth.

Research interests

  • Environmental effects on populations
  • Effects of hunting and predation
  • Time series analysis
  • Spatial population dynamics
  • Variation in demographical parameters
  • Movement and habitat choise
  • Migraiton and phenology
  • Acoustic communication and analysis


Current projects


Read more about my ongoing research and future plans at the Statistical Population Ecology home page.


Scientific publications in peer reviewed journals

Lindroos, L., Engström-Öst, J., Lehtinen, S., Lips, I. & Lindén, A. (in press): Environmental variables driving species and genus level changes in annual plankton biomass. — Journal of Plankton Research, accepted for publication.

Lindén, A., Himberg, M., von Numers, M., Wiklund, T., Engblom, C., Hägerstrand, P., Lill, J.-O. & Hägerstrand, H. (2019): Proportion of river and sea spawning whitefish in catches at the Åland Islands (Baltic Sea), estimated from gill raker counts. — Boreal Environment Research, 24: 101–113.

Otterbeck, A., Vidar Selås, V. Tøttrup Nielsen, J., Roualet, E. & Lindén, A. (2019): The paradox of nest reuse: early breeding benefits reproduction, but nest reuse increases nest predation risk. — Oecologia, 190: 559–568.

Lehikoinen, A., Lindén, A., Karlsson, M., Andersson, A., Crewe, T., Dunn, E., Gregory, G., Karlsson, L., Kristiansen, V., MacKenzie, S., Newman, S., Røer, J. E., Sharpe, C., Sokolov, L., Steinholtz, Å., Stervander, M., Ina, T. & Tjørnløv, R. (2019): Phenology of the avian spring migratory passage in Europe and North America: asymmetric advancement in time and increase in duration. — Ecological Indicators, 101: 985–991.

Forsblom, L., Lehtinen, S. & Lindén, A. (2019): Spatio-temporal population dynamics of six phytoplankton taxa. — Hydrobiologia, 828: 301–314.

Gilljam, D., Knape, J., Lindén, A., Mugabo, M., Sait, S.M. & Fowler, M.S. (2019): The colour of environmental fluctuations associated with terrestrial animal population dynamics. — Global Ecology and Biogeography, 28: 118–130.

Lindén, A. (2018): Adaptive and nonadaptive changes in phenological synchrony. — Proceedings of the National Academy of Sciences of the U.S.A. — 115: 5057–5059.

Öst, M., Lindén, A., Karell, P., Ramula, S. & Kilpi, M. (2018): To breed or not to breed: drivers of intermittent breeding in a seabird under increasing predation risk and male bias. — Oecologia, 188: 129–138.

Ramula, S., Öst, M., Lindén, A., Karell, P. & Kilpi, M. (2018): Increased male bias in eider ducks can be explained by sex-specific survival of prime-age breeders. — PLoS ONE, 13: e0195415.

Papadopoulou, A., Dalsgaard, I., Lindén, A. & Wiklund, T. (2017): In vivo adherence of Flavobacterium psychrophilum to mucosal external surfaces of rainbow trout (Oncorhynchus mykiss) fry. — Journal of Fish Diseases, 40: 1309–1320.

Fraixedas, S., Lindén, A., Meller, K. Lindström, Å., Keišs, O., Kålås, J. A., Husby, M., Leivits, A., Leivits, M. & Lehikoinen, A. (2017): Substantial decline of Northern European peatland bird populations: consequences of drainage. — Biological Conservation, 214: 223–232.

Lindén, A., Meller, K. & Knape, J. (2017): An empirical comparison of models for the phenology of bird migration. — Journal of Avian Biology, 48: 255–265.

Välimäki, K., Lindén, A. & Lehikoinen, A. (2016): Velocity of density shifts in Finnish land bird species depends on their migration ecology and body mass. — Oecologia, 181: 313–321.

Öst, M., Ramula, S., Lindén, A., Karell, P. & Kilpi, M. (2016): Small-scale spatial and temporal variation in the demographic processes underlying he large-scale decline of eiders in the Baltic Sea. — Population Ecology, 58: 121–133.

Fraixedas, S., Lindén, A. & Lehikoinen, A. (2015): Population trends of common breeding forest birds in southern Finland are consistent with trends in forest management and climate change. — Ornis Fennica, 92: 187–203.

Otterbeck, A., Lindén, A. & Roualet, E. (2015): Advantage of specialism: reproductive output is related to prey choice in a small raptor. — Oecologia, 179: 129–137.

Lindén, A., Rapeli, L. & Brutemark, A. (2015): Community attachment and municipal economy: Public attitudes towards wind power in a local context. — Environmental Science & Policy, 54: 10–14.

Ellermaa, M. & Lindén, A. (2015): Sügisränne Põõsaspeal 2014. aastal. [Autumn migration in Cape Põõsaspea in 2014.] — Hirundo, 28: 20–49. (In Estonian with English summary)

Ramula, S., Johansson, J., Lindén, A. & Jonzén, N. (2015): Linking phenological shifts to demographic change. — Climate Research, 63: 135–144.

Fraixedas, S., Lehikoinen, A. & Lindén, A. (2015): Impacts of climate and land-use change on wintering bird populations in Finland. — Journal of Avian Biology, 46: 63–72.

Pakkala, T., Lindén, A., Tiainen, J., Tomppo, E. & Kouki, J. (2014): Indicators of forest biodiversity: which bird species predict breeding bird assemblage diversity in boreal forests at multiple spatial scales? — Annales Zoologici Fennici, 51: 457–476.

Otterbeck, A., Dale, S., Lindén, A. & Marthinsen, G. (2013): A male Reed Warbler and Marsh Warbler hybrid (Acrocephalus scirpaceus × A. palustris) in Norway documented with molecular methods. — Ornis Norvegica, 36: 6–13.

Tornberg, R., Lindén, A., Byholm, P., Ranta, E., Valkama, J., Helle, P. & Lindén, H. (2013): Coupling in goshawk and grouse population dynamics in Finland. — Oecologia, 171: 863–872.

Lehikoinen, A., Lindén. A., Byholm, P., Ranta, E., Saurola, P., Valkama, J., Kaitala, V. & Lindén, H. (2013): Impact of climate change and prey abundance on nesting success of a top-predator, the goshawk. — Oecologia, 171: 283–293.

Lindén, A., Fowler, M. & Jonzén, N. (2013): Mischaracterising density dependence biases estimated effects of coloured covariates on population dynamics. — Population Ecology, 55: 183–192.

Lindén, A. (2011): Using first arrival dates to infer bird migration phenology. — Boreal Environment Research, 16 (Suppl. B): 49–60.

Lehikoinen, A., Lehikoinen, P., Lindén, A. & Laine, T. (2011): Population trend and status of the endangered White-backed Woodpecker Dendrocopos leucotos in Finland. — Ornis Fennica, 88: 195–207.

Knudsen, E., Lindén, A., Both, C., Jonzén, N., Pulido, F., Saino, N., Sutherland, W. J., Bach, L. A., Coppack, T., Ergon, T., Gienapp, P., Gill, J. A., Gordo, O., Hedenström, A., Lehikoinen, E., Marra, P. P., Møller, A. P., Nilsson, A. L. K., Péron, G., Ranta, E., Rubolini, D., Sparks, T. H., Spina, F., Studds, C. E., Sæther, S. A., Tryjanowski, P. & Stenseth, N. C. (2011): Challenging claims in the study of migratory birds and climate change. — Biological Reviews, 86: 928–946.

Lindén, A. & Mäntyniemi, S. (2011): Using negative binomial distribution to model overdispersion in ecological count data. — Ecology, 92: 1414–1421.

Lindén, A., Lehikoinen, A., Hokkanen, T. & Väisänen, R. A. (2011): Modelling irruptions and population dynamics of the great spotted woodpecker — joint effects of density and cone crops. — Oikos, 120: 1065–1075.

Lehikoinen, A., Saurola, P., Byholm, P., Lindén, A. & Valkama, J. (2010): Life history events of Eurasian sparrowhawk in a changing climate. — Journal of Avian Biology, 41: 627–636.

Sirkiä, S., Lindén, A., Helle, P., Nikula, A., Knape, J. & Lindén, H. (2010): Are the declining trends in forest grouse populations due to changes in the forest age structure? A case study of Capercaillie in Finland. — Biological Conservation, 143: 1540–1548.

Ruokolainen, L., Lindén, A., Fowler, M. S. & Kaitala, V. (2009): Ecological and evolutionary dynamics under coloured environmental variation. — Trends in Ecology & Evolution, 24: 555–563.

Lindén, A. & Knape, J. (2009): Estimating environmental effects on population dynamics: consequences of observation error. — Oikos, 118: 675–680.

Knudsen, E., Lindén, A., Ergon, T., Jonzén, N., Vik, J. O., Knape, J., Røer, J. E. & Stenseth, N. C. (2007): Characterizing bird migration phenology using data from standardized monitoring at bird observatories. — Climate Research, 35: 59–77.

Lindholm, A. & Lindén, A. (2007): Some notes on the distribution and songs of two Oriental Cuckoo taxa, Cuculus (saturatus) saturatus and Cuculus (saturatus) optatus. — Forktail, 23: 12–16.

Piha, M., Lindén, A., Pakkala, T. & Tiainen, J. (2007): Linking weather and habitat to population dynamics of a migratory farmland songbird. — Annales Zoologici Fennici, 44: 20–34.

Jonzén, N., Lindén, A., Ergon, T., Knudsen, E., Vik, J. O., Rubolini, D., Piacentini, D., Brinch, C., Spina, F., Karlsson, L., Stervander, M., Andersson, A., Waldenström, J., Lehikoinen, A., Edvardsen, E., Solvang, R. & Stenseth, N. C. (2007): Response to Comment on “Rapid Advance of Spring Arrival Dates in Long-Distance Migratory Birds”. — Science, 315: 598.

Jonzén, N., Lindén, A., Ergon, T., Knudsen, E., Vik, J. O., Rubolini, D., Piacentini, D., Brinch, C., Spina, F., Karlsson, L., Stervander, M., Andersson, A., Waldenström, J., Lehikoinen, A., Edvardsen, E., Solvang, R. & Stenseth, N. C. (2006): Rapid Advance of Spring Arrival Dates in Long-Distance Migratory Birds. — Science, 312: 1959–1961.